Jonathan’s research interests centre around the effects of plants and their root-associated microbial populations on soil properties.  He has developed methods using isotopic tracers of carbon and nutrient elements such as nitrogen and phosphorus, to study the processes by which carbon fixed by photosynthesis in plant shoots is allocated to roots and root-associated symbiotic fungi such as mycorrhizas to mobilize nutrients from soils, and the effects of these  plant-microbial-soil interactions on soil carbon and rates of mineral weathering. His research spans evolutionary biology and understanding how the evolution of plants and their mycorrhizal fungal partners have changed weathering processes and soil development– potentially feeding back to affect global climate via the geochermical carbon cycle.  He is also seeking to understand how additions of calcium silicate minerals to forests and crops can influence plant growth, nutrition, plant health and biogeochemical cycling, helping to sequester carbon from the atmosphere as part of a system to deliver more sustainable agricultural and forestry production.